Browse dbCAN-PUL Entries

PULID Characterization Method(s) Substrate Organism Publication Publish Date Type Num Genes Num CAZymes CazyFamily
PUL0117 expression of recombinant proteins, RNA-seq, differential gene expression, enzyme specificity assay, enzyme activity assay host glycan Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/31160824/>31160824</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/23943617/>23943617</a>
Complex N-glycan breakdown by gut Bacteroides involves an extensive enzymatic apparatus encoded by multiple co-regulated genetic loci. Discovery of beta-1,4-D-mannosyl-N-acetyl-D-glucosamine phosphorylase involved in the metabolism of N-glycans. Nat Microbiol. 2019 Sep;4(9):1571-1581. doi: 10.1038/s41564-019-0466-x. Epub 2019 Jun 3. J Biol Chem. 2013 Sep 20;288(38):27366-27374. doi: 10.1074/jbc.M113.469080. Epub 2013 Aug 13.
2019 Sep,2013 Sep 20 degradation 22 7 GH130_2, GH163, GH18, GH20, GH92
PUL0120 expression of recombinant proteins, RNA-seq, differential gene expression host glycan Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/31160824/>31160824</a>
Complex N-glycan breakdown by gut Bacteroides involves an extensive enzymatic apparatus encoded by multiple co-regulated genetic loci. Nat Microbiol. 2019 Sep;4(9):1571-1581. doi: 10.1038/s41564-019-0466-x. Epub 2019 Jun 3.
2019 Sep degradation 6 1 GH18
PUL0144 enzyme activity assay, Western Blot chitin Thermococcus kodakarensis <a href=https://pubmed.ncbi.nlm.nih.gov/16199574/>16199574</a>
Characterization of a novel glucosamine-6-phosphate deaminase from a hyperthermophilic archaeon. J Bacteriol. 2005 Oct;187(20):7038-44. doi: 10.1128/JB.187.20.7038-7044.2005.
2005 Oct degradation 12 4 CE14, GH1, GH18, GH35
PUL0171 qRT-PCR, RNA-seq host glycan Bacteroides fragilis <a href=https://pubmed.ncbi.nlm.nih.gov/27353652/>27353652</a>
cis-Encoded Small RNAs, a Conserved Mechanism for Repression of Polysaccharide Utilization in Bacteroides. J Bacteriol. 2016 Aug 25;198(18):2410-8. doi: 10.1128/JB.00381-16. Print 2016 Sep 15.
2016 Sep 15 degradation 7 1 GH18
PUL0208 growth assay, clone and expression, enzyme activity assay chitin Pseudoalteromonas luteoviolacea <a href=https://pubmed.ncbi.nlm.nih.gov/31213521/>31213521</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/10220172/>10220172</a>
Marine Chitinolytic Pseudoalteromonas Represents an Untapped Reservoir of Bioactive Potential. Multiple genes involved in chitin degradation from the marine bacterium Pseudoalteromonas sp. strain S91. mSystems. 2019 Jun 18;4(4):e00060-19. doi: 10.1128/mSystems.00060-19. Microbiology (Reading). 1999 Apr;145 ( Pt 4):925-934. doi: 10.1099/13500872-145-4-925.
2019 Jun 18,1999 Apr degradation 3 3 AA10, CBM5, CBM5, GH18, GH18, CBM5, CBM5
PUL0211 enzyme activity assay, gene deletion mutant and growth assay, thin-layer chromatography host glycan Xanthomonas campestris pv. campestris <a href=https://pubmed.ncbi.nlm.nih.gov/25586188/>25586188</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/25205095/>25205095</a>
The N-Glycan cluster from Xanthomonas campestris pv. campestris: a toolbox for sequential plant N-glycan processing. The plant pathogen Xanthomonas campestris pv. campestris exploits N-acetylglucosamine during infection. J Biol Chem. 2015 Mar 6;290(10):6022-36. doi: 10.1074/jbc.M114.624593. Epub 2015 Jan 13. mBio. 2014 Sep 9;5(5):e01527-14. doi: 10.1128/mBio.01527-14.
2015 Mar 6,2014 Sep 9 degradation 9 8 GH125, GH18, GH2, GH20, GH29, GH3, GH35, GH92
PUL0269 RT-PCR chitin Haloferax mediterranei <a href=https://pubmed.ncbi.nlm.nih.gov/23674154/>23674154</a>
Characterization of genes for chitin catabolism in Haloferax mediterranei. Appl Microbiol Biotechnol. 2014 Feb;98(3):1185-94. doi: 10.1007/s00253-013-4969-8. Epub 2013 May 15.
2014 Feb degradation 16 6 CBM5, CBM5, GH18, CE14, GH3
PUL0307 enzyme activity assay chitin Serratia marcescens subsp. marcescens <a href=https://pubmed.ncbi.nlm.nih.gov/29229757/>29229757</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/8757722/>8757722</a>
Structure and activity of ChiX: a peptidoglycan hydrolase required for chitinase secretion by Serratia marcescens. Comparative studies of chitinases A and B from Serratia marcescens. Biochem J. 2018 Jan 23;475(2):415-428. doi: 10.1042/BCJ20170633. Microbiology (Reading). 1996 Jul;142 ( Pt 7):1581-9. doi: 10.1099/13500872-142-7-1581.
2018 Jan 23,1996 Jul degradation 6 2 AA10, GH18
PUL0344 gene deletion mutant and growth assay, protein structure characterization chitin Flavobacterium johnsoniae <a href=https://pubmed.ncbi.nlm.nih.gov/27933102/>27933102</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/32792608/>32792608</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/19717629/>19717629</a>
A polysaccharide utilization locus from Flavobacterium johnsoniae enables conversion of recalcitrant chitin. Structural insights of the enzymes from the chitin utilization locus of Flavobacterium johnsoniae. Novel features of the polysaccharide-digesting gliding bacterium Flavobacterium johnsoniae as revealed by genome sequence analysis. Biotechnol Biofuels. 2016 Nov 28;9:260. doi: 10.1186/s13068-016-0674-z. eCollection 2016. Sci Rep. 2020 Aug 13;10(1):13775. doi: 10.1038/s41598-020-70749-w. Appl Environ Microbiol. 2009 Nov;75(21):6864-75. doi: 10.1128/AEM.01495-09. Epub 2009 Aug 28.
2016,2020 Aug 13,2009 Nov degradation 11 3 GH18, GH18, GH18, GH20
PUL0356 microarray human milk oligosaccharide Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/16968696/>16968696</a>
Functional genomic and metabolic studies of the adaptations of a prominent adult human gut symbiont, Bacteroides thetaiotaomicron, to the suckling period. J Biol Chem. 2006 Nov 24;281(47):36269-79. doi: 10.1074/jbc.M606509200. Epub 2006 Sep 12.
2006 Nov 24 degradation 9 1 GH18
PUL0376 microarray, qPCR mucin Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>
Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007.
2008 Nov 13 degradation 9 2 GH16_3, GH18
PUL0380 microarray, qPCR, microarray, enzyme activity assay, strcutural analysis, clone and expression mucin Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/16968696/>16968696</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/32060313/>32060313</a>
Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Functional genomic and metabolic studies of the adaptations of a prominent adult human gut symbiont, Bacteroides thetaiotaomicron, to the suckling period. Structural basis of mammalian high-mannose N-glycan processing by human gut Bacteroides. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007. J Biol Chem. 2006 Nov 24;281(47):36269-79. doi: 10.1074/jbc.M606509200. Epub 2006 Sep 12. Nat Commun. 2020 Feb 14;11(1):899. doi: 10.1038/s41467-020-14754-7.
2008 Nov 13,2006 Nov 24,2020 Feb 14 degradation 12 4 GH18, GH92
PUL0397 gene deletion mutant and growth assay glycosaminoglycan Capnocytophaga canimorsus <a href=https://pubmed.ncbi.nlm.nih.gov/21762219/>21762219</a>
The genome and surface proteome of Capnocytophaga canimorsus reveal a key role of glycan foraging systems in host glycoproteins deglycosylation. Mol Microbiol. 2011 Aug;81(4):1050-60. doi: 10.1111/j.1365-2958.2011.07750.x. Epub 2011 Jul 18.
2011 Aug degradation 5 1 GH18
PUL0406 high-performance anion-exchange chromatography beta-glucan Coprothermobacter proteolyticus <a href=https://pubmed.ncbi.nlm.nih.gov/30315317/>30315317</a>
From proteins to polysaccharides: lifestyle and genetic evolution of Coprothermobacter proteolyticus. ISME J. 2019 Mar;13(3):603-617. doi: 10.1038/s41396-018-0290-y. Epub 2018 Oct 12.
2019 Mar degradation 21 3 GH16_3, GH18, GH3
PUL0422 microarray host glycan Bacteroides ovatus <a href=https://pubmed.ncbi.nlm.nih.gov/22205877/>22205877</a>
Recognition and degradation of plant cell wall polysaccharides by two human gut symbionts. PLoS Biol. 2011 Dec;9(12):e1001221. doi: 10.1371/journal.pbio.1001221. Epub 2011 Dec 20.
2011 Dec degradation 4 1 GH18
PUL0424 microarray host glycan Bacteroides ovatus <a href=https://pubmed.ncbi.nlm.nih.gov/22205877/>22205877</a>
Recognition and degradation of plant cell wall polysaccharides by two human gut symbionts. PLoS Biol. 2011 Dec;9(12):e1001221. doi: 10.1371/journal.pbio.1001221. Epub 2011 Dec 20.
2011 Dec degradation 18 5 GH130_2, GH163, GH18, GH92
PUL0426 microarray mucin Bacteroides ovatus <a href=https://pubmed.ncbi.nlm.nih.gov/22205877/>22205877</a>
Recognition and degradation of plant cell wall polysaccharides by two human gut symbionts. PLoS Biol. 2011 Dec;9(12):e1001221. doi: 10.1371/journal.pbio.1001221. Epub 2011 Dec 20.
2011 Dec degradation 13 4 GH18, GH92
PUL0442 mass spectrometry, high-performance anion-exchange chromatography, SDS-PAGE, recombinant protein expression, enzyme activity assay, substrate binding assay glucomannan/chitin Chitinophaga pinensis DSM 2588 <a href=https://pubmed.ncbi.nlm.nih.gov/28069559/>28069559</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/37493618/>37493618</a>
Proteomic insights into mannan degradation and protein secretion by the forest floor bacterium Chitinophaga pinensis. A polysaccharide utilization locus from Chitinophaga pinensis simultaneously targets chitin and beta-glucans found in fungal cell walls. J Proteomics. 2017 Mar 6;156:63-74. doi: 10.1016/j.jprot.2017.01.003. Epub 2017 Jan 6. mSphere. 2023 Aug 24;8(4):e0024423. doi: 10.1128/msphere.00244-23. Epub 2023 Jul 26.
2017 Mar 6,2023 Aug 24 degradation 9 3 GH16_3, CBM6, GH18, CBM6, GH18, GH18, CBM5
PUL0465 microarray, qPCR mucin Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>
Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007.
2008 Nov 13 degradation 8 2 GH18
PUL0467 microarray, qPCR, expression of recombinant proteins, RNA-seq, differential gene expression host glycan Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/31160824/>31160824</a>
Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Complex N-glycan breakdown by gut Bacteroides involves an extensive enzymatic apparatus encoded by multiple co-regulated genetic loci. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007. Nat Microbiol. 2019 Sep;4(9):1571-1581. doi: 10.1038/s41564-019-0466-x. Epub 2019 Jun 3.
2008 Nov 13,2019 Sep degradation 14 4 GH18, GH2, GH20, GH29, CBM32
PUL0470 microarray, qPCR mucin Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>
Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007.
2008 Nov 13 degradation 7 1 GH18
PUL0475 clone and expression, gene deletion mutant and growth assay beta-glucan Streptomyces reticuli <a href=https://pubmed.ncbi.nlm.nih.gov/10347054/>10347054</a>
Characterization of the binding protein-dependent cellobiose and cellotriose transport system of the cellulose degrader Streptomyces reticuli. Appl Environ Microbiol. 1999 Jun;65(6):2636-43. doi: 10.1128/AEM.65.6.2636-2643.1999.
1999 Jun degradation 7 2 CBM2, GH18, GH1
PUL0497 clone and expression, enzyme activity assay chitin Pseudoalteromonas piscicida <a href=https://pubmed.ncbi.nlm.nih.gov/11772635/>11772635</a>
Identification and characterization of the gene cluster involved in chitin degradation in a marine bacterium, Alteromonas sp. strain O-7. Appl Environ Microbiol. 2002 Jan;68(1):263-70. doi: 10.1128/AEM.68.1.263-270.2002.
2002 Jan degradation 3 3 AA10, CBM5, GH18, GH18, CBM5
PUL0556 gene deletion mutant and growth assay, qRT-PCR, microarray, enzyme activity assay host glycan Bacteroides fragilis <a href=https://pubmed.ncbi.nlm.nih.gov/25139987/>25139987</a>
Efficient utilization of complex N-linked glycans is a selective advantage for Bacteroides fragilis in extraintestinal infections. Proc Natl Acad Sci U S A. 2014 Sep 2;111(35):12901-6. doi: 10.1073/pnas.1407344111. Epub 2014 Aug 19.
2014 Sep 2 degradation 8 2 GH18, GH97
PUL0665 UHPLC-MS, RNA-seq, RT-qPCR pectin Bacteroides thetaiotaomicron VPI-5482 <a href=https://pubmed.ncbi.nlm.nih.gov/34420703/>34420703</a>
Discrete genetic loci in human gut Bacteroides thetaiotaomicron confer pectin metabolism. Carbohydr Polym. 2021 Nov 15;272:118534. doi: 10.1016/j.carbpol.2021.118534. Epub 2021 Aug 6.
2021 Nov 15 degradation 8 2 GH18, GH30_4
PUL0786 RNA-seq, reducing-sugar assay, growth assay, high performance gel permeation chromatography, gas chromatography, RNA-seq, differential gene expression pectic polysaccharide Bacteroides ovatus strain ATCC 8483 <a href=https://pubmed.ncbi.nlm.nih.gov/38890895/>38890895</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/39892338/>39892338</a>
The Utilization by Bacteroides spp. of a Purified Polysaccharide from Fuzhuan Brick Tea. In vitro fermentation of a purified fraction of polysaccharides from the root of Brassica rapa L. by human gut microbiota and its interaction with Bacteroides ovatus. Foods. 2024 May 26;13(11):1666. doi: 10.3390/foods13111666. Food Chem. 2025 May 1;473:143109. doi: 10.1016/j.foodchem.2025.143109. Epub 2025 Jan 27.
2024 May 26,2025 May 1 degradation 26 14 CBM67, GH78, CBM67, GH78, GH33, CE19, GH105, GH130_2, GH140, GH143, GH142, GH163, GH18, GH28, GH43_18, GH92, GH95, PL1_2