Browse dbCAN-PUL Entries

PULID Characterization Method(s) Substrate Organism Publication Publish Date Type Num Genes Num CAZymes CazyFamily
PUL0031 RNA-seq starch Bifidobacterium longum <a href=https://pubmed.ncbi.nlm.nih.gov/16523284/>16523284</a>
A functional analysis of the Bifidobacterium longum cscA and scrP genes in sucrose utilization. Appl Microbiol Biotechnol. 2006 Oct;72(5):975-81. doi: 10.1007/s00253-006-0358-x. Epub 2006 Mar 8.
2006 Oct degradation 3 1 GH32
PUL0032 RNA-seq starch Bifidobacterium longum <a href=https://pubmed.ncbi.nlm.nih.gov/16523284/>16523284</a>
A functional analysis of the Bifidobacterium longum cscA and scrP genes in sucrose utilization. Appl Microbiol Biotechnol. 2006 Oct;72(5):975-81. doi: 10.1007/s00253-006-0358-x. Epub 2006 Mar 8.
2006 Oct degradation 3 1 GH13_18
PUL0042 RT-PCR starch Caulobacter vibrioides <a href=https://pubmed.ncbi.nlm.nih.gov/30054816/>30054816</a>
SucA-dependent uptake of sucrose across the outer membrane of Caulobacter crescentus. J Microbiol. 2018 Sep;56(9):648-655. doi: 10.1007/s12275-018-8225-x. Epub 2018 Jul 27.
2018 Sep degradation 5 1 GH13_4
PUL0141 sequence homology analysis starch Bifidobacterium animalis subsp. animalis <a href=https://pubmed.ncbi.nlm.nih.gov/30306201/>30306201</a>
Staying alive: growth and survival of Bifidobacterium animalis subsp. animalis under in vitro and in vivo conditions. Appl Microbiol Biotechnol. 2018 Dec;102(24):10645-10663. doi: 10.1007/s00253-018-9413-7. Epub 2018 Oct 10.
2018 Dec degradation 4 1 GH13_18
PUL0142 sequence homology analysis starch Bifidobacterium animalis subsp. animalis <a href=https://pubmed.ncbi.nlm.nih.gov/30306201/>30306201</a>
Staying alive: growth and survival of Bifidobacterium animalis subsp. animalis under in vitro and in vivo conditions. Appl Microbiol Biotechnol. 2018 Dec;102(24):10645-10663. doi: 10.1007/s00253-018-9413-7. Epub 2018 Oct 10.
2018 Dec degradation 5 1 GH13_30
PUL0166 enzyme activity assay, RT-PCR starch Bacteroides fragilis <a href=https://pubmed.ncbi.nlm.nih.gov/16788175/>16788175</a>
Characterization of the primary starch utilization operon in the obligate anaerobe Bacteroides fragilis: Regulation by carbon source and oxygen. J Bacteriol. 2006 Jul;188(13):4663-72. doi: 10.1128/JB.00125-06.
2006 Jul degradation 5 1 GH13_10
PUL0174 RT-PCR, enzyme activity assay, enzymatic product analysis starch Kribbella flavida <a href=https://pubmed.ncbi.nlm.nih.gov/27302067/>27302067</a>
Two Novel Glycoside Hydrolases Responsible for the Catabolism of Cyclobis-(1-->6)-alpha-nigerosyl. J Biol Chem. 2016 Aug 5;291(32):16438-47. doi: 10.1074/jbc.M116.727305. Epub 2016 Jun 14.
2016 Aug 5 degradation 3 2 GH31_12, CBM20, GH31_7, CBM35
PUL0176 RT-PCR, enzyme activity assay, enzymatic product analysis starch Kribbella flavida <a href=https://pubmed.ncbi.nlm.nih.gov/27302067/>27302067</a>
Two Novel Glycoside Hydrolases Responsible for the Catabolism of Cyclobis-(1-->6)-alpha-nigerosyl. J Biol Chem. 2016 Aug 5;291(32):16438-47. doi: 10.1074/jbc.M116.727305. Epub 2016 Jun 14.
2016 Aug 5 degradation 6 2 GH15, GH31_7
PUL0197 gene deletion mutant and growth assay starch Streptococcus mutans <a href=https://pubmed.ncbi.nlm.nih.gov/17233733/>17233733</a>
Overlapping substrate specificity for sucrose and maltose of two binding protein-dependent sugar uptake systems in Streptococcus mutans. FEMS Microbiol Lett. 2007 Jan;266(2):218-23. doi: 10.1111/j.1574-6968.2006.00522.x.
2007 Jan degradation 7 2 GH77, GT35
PUL0204 qPCR, thin-layer chromatography, substrate binding assay starch Bacteroides thetaiotaomicron <a href=https://pubmed.ncbi.nlm.nih.gov/25841008/>25841008</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/18996345/>18996345</a>
Differential Metabolism of Exopolysaccharides from Probiotic Lactobacilli by the Human Gut Symbiont Bacteroides thetaiotaomicron. Mucosal glycan foraging enhances fitness and transmission of a saccharolytic human gut bacterial symbiont. Appl Environ Microbiol. 2015 Jun 15;81(12):3973-83. doi: 10.1128/AEM.00149-15. Epub 2015 Apr 3. Cell Host Microbe. 2008 Nov 13;4(5):447-57. doi: 10.1016/j.chom.2008.09.007.
2015 Jun 15,2008 Nov 13 degradation 7 3 GH13_36, GH13_46, GH97
PUL0230 RT-PCR, enzyme activity assay, clone, enzyme kinetic analysis, thin-layer chromatography, crystallization starch Lactobacillus acidophilus <a href=https://pubmed.ncbi.nlm.nih.gov/22685275/>22685275</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/32444471/>32444471</a>
Enzymology and structure of the GH13_31 glucan 1,6-alpha-glucosidase that confers isomaltooligosaccharide utilization in the probiotic Lactobacillus acidophilus NCFM. An 1,4-alpha-Glucosyltransferase Defines a New Maltodextrin Catabolism Scheme in Lactobacillus acidophilus. J Bacteriol. 2012 Aug;194(16):4249-59. doi: 10.1128/JB.00622-12. Epub 2012 Jun 8. Appl Environ Microbiol. 2020 Jul 20;86(15):e00661-20. doi: 10.1128/AEM.00661-20. Print 2020 Jul 20.
2012 Aug,2020 Jul 20 degradation 12 3 CBM34, GH13_20, GH13_31, GH65
PUL0265 enzyme activity assay, gene deletion mutant and growth assay starch Staphylococcus xylosus <a href=https://pubmed.ncbi.nlm.nih.gov/7730272/>7730272</a>
Characterization of a genetic locus essential for maltose-maltotriose utilization in Staphylococcus xylosus. J Bacteriol. 1995 May;177(9):2408-15. doi: 10.1128/jb.177.9.2408-2415.1995.
1995 May degradation 2 1 GH13_31
PUL0268 Northern Blot, promoter assay starch Geobacillus kaustophilus <a href=https://pubmed.ncbi.nlm.nih.gov/23793634/>23793634</a>
Polysaccharide-degrading thermophiles generated by heterologous gene expression in Geobacillus kaustophilus HTA426. Appl Environ Microbiol. 2013 Sep;79(17):5151-8. doi: 10.1128/AEM.01506-13. Epub 2013 Jun 21.
2013 Sep degradation 5 1 GH13_45
PUL0275 RT-qPCR starch Bifidobacterium animalis subsp. lactis <a href=https://pubmed.ncbi.nlm.nih.gov/23663691/>23663691</a>
Transcriptional analysis of oligosaccharide utilization by Bifidobacterium lactis Bl-04. BMC Genomics. 2013 May 10;14:312. doi: 10.1186/1471-2164-14-312.
2013 May 10 degradation 8 3 GH13_30, GH13_44, GH77
PUL0276 RT-qPCR starch Bifidobacterium animalis subsp. lactis <a href=https://pubmed.ncbi.nlm.nih.gov/23663691/>23663691</a>
Transcriptional analysis of oligosaccharide utilization by Bifidobacterium lactis Bl-04. BMC Genomics. 2013 May 10;14:312. doi: 10.1186/1471-2164-14-312.
2013 May 10 degradation 9 3 GH13_31, GH36
PUL0312 RT-PCR starch Gluconacetobacter diazotrophicus <a href=https://pubmed.ncbi.nlm.nih.gov/19139238/>19139238</a>
Transcriptional regulation and signal-peptide-dependent secretion of exolevanase (LsdB) in the endophyte Gluconacetobacter diazotrophicus. Appl Environ Microbiol. 2009 Mar;75(6):1782-5. doi: 10.1128/AEM.01887-08. Epub 2009 Jan 9.
2009 Mar degradation 2 2 GH32, GH68
PUL0327 microarray, gas chromatography, mass spectrometry, gene deletion mutant and growth assay, complementation study starch Enterococcus faecalis <a href=https://pubmed.ncbi.nlm.nih.gov/28455338/>28455338</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/32680872/>32680872</a>
Enzymes Required for Maltodextrin Catabolism in Enterococcus faecalis Exhibit Novel Activities. Enterococcus faecalis Maltodextrin Gene Regulation by Combined Action of Maltose Gene Regulator MalR and Pleiotropic Regulator CcpA. Appl Environ Microbiol. 2017 Jun 16;83(13):e00038-17. doi: 10.1128/AEM.00038-17. Print 2017 Jul 1. Appl Environ Microbiol. 2020 Sep 1;86(18):e01147-20. doi: 10.1128/AEM.01147-20. Print 2020 Sep 1.
2017 Jul 1,2020 Sep 1 degradation 6 2 CBM34, GH13_20, GH13_31
PUL0347 gene deletion mutant and growth assay, RT-PCR starch Bifidobacterium breve <a href=https://pubmed.ncbi.nlm.nih.gov/24581150/>24581150</a>
Comparative genomics of the Bifidobacterium breve taxon. BMC Genomics. 2014 Mar 1;15(1):170. doi: 10.1186/1471-2164-15-170.
2014 Mar 1 degradation 13 1 GH13_11
PUL0349 microarray starch Leptotrichia buccalis <a href=https://pubmed.ncbi.nlm.nih.gov/22230464/>22230464</a>
Metabolism of sugars by genetically diverse species of oral Leptotrichia. Mol Oral Microbiol. 2012 Feb;27(1):34-44. doi: 10.1111/j.2041-1014.2011.00627.x. Epub 2011 Oct 4.
2012 Feb degradation 3 1 GH4
PUL0351 enzyme activity assay starch Escherichia coli <a href=https://pubmed.ncbi.nlm.nih.gov/1435727/>1435727</a>
Characterization of a chromosomally encoded, non-PTS metabolic pathway for sucrose utilization in Escherichia coli EC3132. Mol Gen Genet. 1992 Oct;235(1):22-32. doi: 10.1007/BF00286177.
1992 Oct degradation 4 1 GH32
PUL0361 enzyme activity assay starch Thermotoga maritima <a href=https://pubmed.ncbi.nlm.nih.gov/10972187/>10972187</a>
Thermotoga maritima AglA, an extremely thermostable NAD+-, Mn2+-, and thiol-dependent alpha-glucosidase. Extremophiles. 2000 Aug;4(4):189-200. doi: 10.1007/pl00010711.
2000 Aug degradation 6 3 GH13_20, GH13_36, GH4
PUL0362 enzyme activity assay starch Xanthomonas campestris pv. campestris <a href=https://pubmed.ncbi.nlm.nih.gov/17311090/>17311090</a>
Plant carbohydrate scavenging through tonB-dependent receptors: a feature shared by phytopathogenic and aquatic bacteria. PLoS One. 2007 Feb 21;2(2):e224. doi: 10.1371/journal.pone.0000224.
2007 Feb 21 degradation 4 1 GH13_4
PUL0365 RT-PCR starch Xanthomonas campestris pv. campestris <a href=https://pubmed.ncbi.nlm.nih.gov/17311090/>17311090</a>
Plant carbohydrate scavenging through tonB-dependent receptors: a feature shared by phytopathogenic and aquatic bacteria. PLoS One. 2007 Feb 21;2(2):e224. doi: 10.1371/journal.pone.0000224.
2007 Feb 21 degradation 8 4 GH13_2, GH13_23, GH78, GH97
PUL0371 enzyme activity assay starch Thermococcus sp. B1001 <a href=https://pubmed.ncbi.nlm.nih.gov/11489857/>11489857</a>
Extracellular synthesis, specific recognition, and intracellular degradation of cyclomaltodextrins by the hyperthermophilic archaeon Thermococcus sp. strain B1001. J Bacteriol. 2001 Sep;183(17):5050-7. doi: 10.1128/JB.183.17.5050-5057.2001.
2001 Sep degradation 5 2 CBM34, GH13_20, GH13_2, CBM20
PUL0373 enzyme activity assay starch Dickeya dadantii <a href=https://pubmed.ncbi.nlm.nih.gov/19734309/>19734309</a>
Catabolism of raffinose, sucrose, and melibiose in Erwinia chrysanthemi 3937. J Bacteriol. 2009 Nov;191(22):6960-7. doi: 10.1128/JB.00594-09. Epub 2009 Sep 4.
2009 Nov degradation 5 1 GH32
PUL0421 microarray starch Bacteroides ovatus <a href=https://pubmed.ncbi.nlm.nih.gov/22205877/>22205877</a>
Recognition and degradation of plant cell wall polysaccharides by two human gut symbionts. PLoS Biol. 2011 Dec;9(12):e1001221. doi: 10.1371/journal.pbio.1001221. Epub 2011 Dec 20.
2011 Dec degradation 7 3 GH13_10, GH13_46, GH97
PUL0455 clone and expression, genes induced in presence of substrate, enzyme activity assay starch Bifidobacterium animalis <a href=https://pubmed.ncbi.nlm.nih.gov/12513973/>12513973</a>
Induction of sucrose utilization genes from Bifidobacterium lactis by sucrose and raffinose. Appl Environ Microbiol. 2003 Jan;69(1):24-32. doi: 10.1128/AEM.69.1.24-32.2003.
2003 Jan degradation 3 1 GH13_18
PUL0485 growth assay, qRT-PCR, enzyme activity assay, affinity gel electrophoresis, crystallization, recombinant protein expression starch Flavobacterium johnsoniae <a href=https://pubmed.ncbi.nlm.nih.gov/19717629/>19717629</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/37269952/>37269952</a>, <a href=https://pubmed.ncbi.nlm.nih.gov/38661728/>38661728</a>
Novel features of the polysaccharide-digesting gliding bacterium Flavobacterium johnsoniae as revealed by genome sequence analysis. Bacteroidota polysaccharide utilization system for branched dextran exopolysaccharides from lactic acid bacteria. Structural insights into alpha-(1-->6)-linkage preference of GH97 glucodextranase from Flavobacterium johnsoniae. Appl Environ Microbiol. 2009 Nov;75(21):6864-75. doi: 10.1128/AEM.01495-09. Epub 2009 Aug 28. J Biol Chem. 2023 Jul;299(7):104885. doi: 10.1016/j.jbc.2023.104885. Epub 2023 Jun 2. FEBS J. 2024 Jul;291(14):3267-3282. doi: 10.1111/febs.17139. Epub 2024 Apr 25.
2009 Nov,2023 Jul,2024 Jul degradation 9 5 GH27, CBM13, GH31, GH65, GH66, GH97
PUL0519 gene deletion mutant and growth assay starch Streptococcus pneumoniae <a href=https://pubmed.ncbi.nlm.nih.gov/8244973/>8244973</a>
Characterization of the Streptococcus pneumoniae maltosaccharide regulator MalR, a member of the LacI-GalR family of repressors displaying distinctive genetic features. J Biol Chem. 1993 Dec 5;268(34):25402-8.
1993 Dec 5 degradation 7 2 GH77, GT35
PUL0540 RNA-seq starch Bacteroides cellulosilyticus <a href=https://pubmed.ncbi.nlm.nih.gov/23976882/>23976882</a>
Effects of diet on resource utilization by a model human gut microbiota containing Bacteroides cellulosilyticus WH2, a symbiont with an extensive glycobiome. PLoS Biol. 2013;11(8):e1001637. doi: 10.1371/journal.pbio.1001637. Epub 2013 Aug 20.
2013 degradation 6 3 GH13, GH97
PUL0554 RNA-seq starch Bacteroides cellulosilyticus <a href=https://pubmed.ncbi.nlm.nih.gov/23976882/>23976882</a>
Effects of diet on resource utilization by a model human gut microbiota containing Bacteroides cellulosilyticus WH2, a symbiont with an extensive glycobiome. PLoS Biol. 2013;11(8):e1001637. doi: 10.1371/journal.pbio.1001637. Epub 2013 Aug 20.
2013 degradation 6 3 GH31, GH66, GH97
PUL0568 clone and expression, enzyme activity assay, Northern Blot starch Clostridium beijerinckii <a href=https://pubmed.ncbi.nlm.nih.gov/10411273/>10411273</a>
The genes controlling sucrose utilization in Clostridium beijerinckii NCIMB 8052 constitute an operon. Microbiology (Reading). 1999 Jun;145 ( Pt 6):1461-1472. doi: 10.1099/13500872-145-6-1461.
1999 Jun degradation 4 1 GH32
PUL0589 Western Blot, enzyme activity assay, thin-layer chromatography starch Streptococcus mutans <a href=https://pubmed.ncbi.nlm.nih.gov/23930155/>23930155</a>
The malQ gene is essential for starch metabolism in Streptococcus mutans. J Oral Microbiol. 2013 Aug 6;5. doi: 10.3402/jom.v5i0.21285. Print 2013.
2013 degradation 3 2 GH77, GT35
PUL0590 qRT-PCR, gene deletion mutant and growth assay, microarray starch Enterococcus faecium <a href=https://pubmed.ncbi.nlm.nih.gov/23951303/>23951303</a>
A LacI-family regulator activates maltodextrin metabolism of Enterococcus faecium. PLoS One. 2013 Aug 7;8(8):e72285. doi: 10.1371/journal.pone.0072285. eCollection 2013.
2013 degradation 5 1 CBM34, GH13_20
PUL0593 Northern Blot starch Thermoanaerobacterium thermosulfurigenes <a href=https://pubmed.ncbi.nlm.nih.gov/8576036/>8576036</a>
Molecular analysis of the amy gene locus of Thermoanaerobacterium thermosulfurigenes EM1 encoding starch-degrading enzymes and a binding protein-dependent maltose transport system. J Bacteriol. 1996 Feb;178(4):1039-46. doi: 10.1128/jb.178.4.1039-1046.1996.
1996 Feb degradation 6 2 CBM34, GH13_39, CBM20, GH13_2, CBM20
PUL0595 enzyme activity assay, qPCR, crystallization starch [Eubacterium] rectale <a href=https://pubmed.ncbi.nlm.nih.gov/25388295/>25388295</a>
Molecular details of a starch utilization pathway in the human gut symbiont Eubacterium rectale. Mol Microbiol. 2015 Jan;95(2):209-30. doi: 10.1111/mmi.12859. Epub 2014 Dec 19.
2015 Jan degradation 4 1 CBM82, CBM83, GH13_41
PUL0596 enzyme activity assay, qPCR, crystallization starch [Eubacterium] rectale <a href=https://pubmed.ncbi.nlm.nih.gov/25388295/>25388295</a>
Molecular details of a starch utilization pathway in the human gut symbiont Eubacterium rectale. Mol Microbiol. 2015 Jan;95(2):209-30. doi: 10.1111/mmi.12859. Epub 2014 Dec 19.
2015 Jan degradation 4 1 GH13_36
PUL0642 recombinant protein expression starch Roseburia intestinalis L1-82 <a href=https://pubmed.ncbi.nlm.nih.gov/33439065/>33439065</a>
Characterization of fructooligosaccharide metabolism and fructooligosaccharide-degrading enzymes in human commensal butyrate producers. Gut Microbes. 2021 Jan-Dec;13(1):1-20. doi: 10.1080/19490976.2020.1869503.
2021 Jan-Dec degradation 11 6 CBM61, GH53, CBM86, CBM22, GH10, CBM9, GH13_18, GH13_31, GH32, GH36
PUL0647 qPCR starch Streptococcus mutans UA159 <a href=https://pubmed.ncbi.nlm.nih.gov/33603728/>33603728</a>
The Route of Sucrose Utilization by Streptococcus mutans Affects Intracellular Polysaccharide Metabolism. Front Microbiol. 2021 Feb 2;12:636684. doi: 10.3389/fmicb.2021.636684. eCollection 2021.
2021 biosynthesis 5 3 CBM48, GH13_9, GT35, GT5
PUL0714 clone and expression, crystallization, isothermal titration calorimetry (ITC), thin-layer chromatography, Western Blot, gene mutant starch Bacteroides ovatus strain ATCC 8483  <a href=https://pubmed.ncbi.nlm.nih.gov/37500984/>37500984</a>
BoGH13A(Sus) from Bacteroides ovatus represents a novel alpha-amylase used for Bacteroides starch breakdown in the human gut. Cell Mol Life Sci. 2023 Jul 28;80(8):232. doi: 10.1007/s00018-023-04812-w.
2023 Jul 28 degradation 8 3 GH13_10, GH13_46, GH97
PUL0719 RNA-seq, growth assay starch Xanthomonas citri pv. citri str. 306 <a href=https://pubmed.ncbi.nlm.nih.gov/37855631/>37855631</a>
Plant structural and storage glucans trigger distinct transcriptional responses that modulate the motility of Xanthomonas pathogens. Microbiol Spectr. 2023 Dec 12;11(6):e0228023. doi: 10.1128/spectrum.02280-23. Epub 2023 Oct 19.
2023 Dec 12 degradation 7 3 GH13_2, GH13_23, GH97
PUL0766 RNA-seq, qRT-PCR, gas chromatography, mass spectrometry, thin-layer chromatography starch Segatella copri DSM 18205 <a href=https://pubmed.ncbi.nlm.nih.gov/39636128/>39636128</a>
Transcriptional delineation of polysaccharide utilization loci in the human gut commensal Segatella copri DSM18205 and co-culture with exemplar Bacteroides species on dietary plant glycans. Appl Environ Microbiol. 2025 Jan 31;91(1):e0175924. doi: 10.1128/aem.01759-24. Epub 2024 Dec 5.
2025 Jan 31 degradation 5 2 GH13_44, GH97